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5 ), including the ribosomal protein S6 kinase (S6K) and eukaryotic translation initiation factor 4E-binding protein 1 (4E-BP1) that mediate mTORC1 stimulation of protein synthesis 1 , whereas it is dispensable for the phosphorylation of noncanonical substrates, such as transcription factor EB (TFEB) 6

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The first one involves irreversible damage to the ETC, possibly through TTC5 upregulation, resulting in ATP loss, increased mitochondrial superoxide production and caspases 3/7 activation, and subsequent apoptosis