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2016;11:e0165267

Alterations in regulation of energy homeostasis in cyclic nucleotide phosphodiesterase 3Bnull mice

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The imidazole nitrogen of the histidine side chain donates one of the three nitrogen ligands to the copper, and the position of histidine at the second residue is what enables the deprotonated amide nitrogen of the preceding Gly-His peptide bond to also coordinate the copper